Audit Committee Position

John P. Brancaccio

Independent Director since 2008

Chair

Melvin K. Spigelman, M.D.

Independent Director since 2008

Member

Timothy S. Callahan

Independent Director since 2015

Member

Audit Committee Charter

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Compensation Committee Position

Thomas H. Adams, Ph.D.

Independent Director since 2008

Chair

John P. Brancaccio

Independent Director since 2008

Member

Melvin K. Spigelman, M.D.

Independent Director since 2008

Member

Richard J. Daly

Independent Director since 2015

Member

Compensation Committee Charter

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Governance and Nominating Committee Position

John P. Brancaccio

Independent Director since 2008

Chair

Thomas H. Adams, Ph.D.

Independent Director since 2008

Member

Governance and Nominating Committee Charter

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Compliance Committee Position

Timothy S. Callahan

Independent Director since 2015

Chair

Gary S. Jacob, Ph.D.

Chairman of the Board and CEO

Member

John P. Brancaccio

Independent Director since 2008

Member

Alan F. Joslyn, Ph.D.

Director since 2008

Member

Compliance Committee Charter

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