Audit Committee Position

John P. Brancaccio

Independent Director since 2008

Chair

Melvin K. Spigelman, M.D.

Independent Director since 2008

Member

Timothy S. Callahan

Independent Director since 2015

Member

Audit Committee Charter

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Compensation Committee Position

Thomas H. Adams, Ph.D.

Independent Director since 2008

Chair

John P. Brancaccio

Independent Director since 2008

Member

Melvin K. Spigelman, M.D.

Independent Director since 2008

Member

Richard J. Daly

Independent Director since 2015

Member

Compensation Committee Charter

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Governance and Nominating Committee Position

John P. Brancaccio

Independent Director since 2008

Chair

Thomas H. Adams, Ph.D.

Independent Director since 2008

Member

Governance and Nominating Committee Charter

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External Communication Oversight Committee Position

Melvin K. Spigelman, M.D.

Independent Director since 2008

Chair

Alan F. Joslyn, Ph.D.

Director since 2008

Member